Set two years after the defeat of Clockwerk in Sly Cooper and the Thievius Raccoonus, the game begins as Sly and the gang attempt to steal the Clockwerk parts from a Natural History Museum in Cairo, Egypt. It turns out, however, that the parts have been already stolen, and Carmelita Fox along with her new partner Constable Neyla turn up at the scene instead. Constable Neyla purposefully hints that the parts could have been stolen by a group of outlaws known as The Klaww Gang, each member using a Clockwerk body part to aid his or her own personal conquest. Sly and the gang set out to find the missing parts, and destroy them.
The trio first travel to Paris and steal the Clockwerk Tail Feathers back from Dimitri, a club owner and "underworld celebrity" who was using the feathers as a printing press for forged money due their unique alloy being unaffected to wear. After putting the forger behind bars, the gang headed off to India where they steal back the Clockwerk Wings back from Rajan, a tiger who controls the Klaww Gang's illegal spice operations and uses the wings to earn prestige. After pursuing Rajan into the jungle, the trio manage to steal back the Clockwerk Heart. Unfortunately, the gang are betrayed by Neyla who throws them (along with a framed Carmelita) into the care of the Contessa, a high ranking prison warden whose mastery of hypnosis to "re-educate" criminals has earned her a reputation amongst Interpol.
Being the only one not to be captured, Bentley travels to Prague to help bust Sly and Murray from the Contessa's prison. In the process, he learns that she too is a Klaww Gang member, using the illegal spice as part of her hypnosis techniques so that the thieves in her care confess where they have hidden all of their wealth. Though the Contesssa initially escapes, the gang track her down and steal the Clockwerk Eyes from the Contessa, freeing Carmelita in the process. The prison warden is brough to trial for the crimes she commited and Carmelita reluctantly takes Sly's help in an escape (still being a framed cop). The gang then track down Jean Bison, another member of the Klaww Gang who is a shipping baron who owns half the traines in Canada, and steal back the Clockwerk Talons, Stomach and Lungs from him (he was using the talons to clear-cut the forests and the lungs and stomach as bellows that would never wear out in his trains). However, Bison captures the 3 of them in their 2nd adventure on his turf, and then sells all of their hard-earned Clockwerk parts to another Klaww Gang member. The three defeat him and then sneak aboard a battery as part of a Trojan-Horse-type plan which takes them to the buyer's airship. Tragically, they have to leave the van behind, much to Murray's dismay.
The gang's journey eventually culminates in a confrontation with the gang's leader, Arpeggio, on his blimp as well as his protege who is revealed to be Neyla; both have managed to gather all of the parts together to create the Clockwerk Frame. The Klaww Gang leader reveals that he plans to use the parts to become super strong and eventually use the power of hatred Clockwerk used to live for hundreds of years to become immortal. Dimitri is revealed to have gotten all of Paris to consume spice he aqquired from Rajan (who shipped it through Jean Bison) and using the theory of the Contessa's mastery of hypnosis, Arpeggio plans to use the Northen Lights as a giant "Hypno-Light" to fill the people below with hatred. This would render him immortal.
Suddenly, Neyla double-crosses him and bonds with the Clockwerk parts herself. She kills Arpeggio with her beak and proclaims that "Clock-La" is born before flying away. Knowing they need to act fast, the gang disable the blimp's engines to stop Clock-La from drawing energy, After some help from Carmelita, the mechanical hybrid is beaten and crashes down just outside of Paris. After the battle, Clock-La begins to scream at Sly, shouting that his life will never be the safe as long as she is alive, a hint that Clockwerk's real mind is starting to take control.
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